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Harpalus {genus} -
Arthropoda
;
Insecta
;
Coleoptera
;
Carabidae
;
Harpalinae
;
Harpalini
;
Print
Search Taxonomy
Sub-taxa
Species (122)
Harpalus D3_D [1]
Harpalus actiosus [1]
Harpalus aesculanus [1]
Harpalus affinis [119]
Harpalus albanicus [4]
Harpalus amputatus [70]
Harpalus animosus [51]
Harpalus anxius [34]
Harpalus atratus [33]
Harpalus atrichatus [17]
Harpalus attenuatus [15]
Harpalus aureus [1]
Harpalus autumnalis [21]
Harpalus bellieri [5]
Harpalus bungii [2]
Harpalus calceatus [40]
Harpalus caliginosus [147]
Harpalus capito [10]
Harpalus caspius [24]
Harpalus cautus [50]
Harpalus cephalotes [1]
Harpalus cohni [6]
Harpalus compar [79]
Harpalus contemptus [2]
Harpalus cordatus [35]
Harpalus cordifer [29]
Harpalus cupreus [4]
Harpalus decipiens [2]
Harpalus desertus [114]
Harpalus dimidiatus [22]
Harpalus discrepans [1]
Harpalus dispar [2]
Harpalus distinguendus [61]
Harpalus ebeninus [1]
Harpalus ellipsis [125]
Harpalus eous [8]
Harpalus erraticus [22]
Harpalus erythropus [18]
Harpalus faunus [24]
Harpalus flavescens [15]
Harpalus flavicornis [7]
Harpalus fraternus [29]
Harpalus froelichii [41]
Harpalus fulvilabris [54]
Harpalus fuscipalpis [34]
Harpalus gravis [2]
Harpalus griseus [76]
Harpalus hatchi [3]
Harpalus herbivagus [94]
Harpalus hirtipes [7]
Harpalus honestus [30]
Harpalus indianus [10]
Harpalus indigens [57]
Harpalus innocuus [6]
Harpalus jureceki [3]
Harpalus karamani [1]
Harpalus katiae [24]
Harpalus laevipes [35]
Harpalus laticeps [42]
Harpalus latus [66]
Harpalus lewisii [3]
Harpalus luteicornis [13]
Harpalus marginellus [7]
Harpalus megacephalus [4]
Harpalus melancholicus [13]
Harpalus microthorax [2]
Harpalus modestus [3]
Harpalus neglectus [16]
Harpalus nigritarsis [98]
Harpalus niigatanus [2]
Harpalus obliquus [15]
Harpalus oblitus [2]
Harpalus obnixus [5]
Harpalus ochropus [2]
Harpalus opacipennis [133]
Harpalus paratus [90]
Harpalus pensylvanicus [268]
Harpalus picipennis [6]
Harpalus pigmaeus [1]
Harpalus plenalis [3]
Harpalus politus [3]
Harpalus progrediens [3]
Harpalus protractus [41]
Harpalus providens [78]
Harpalus pumilus [32]
Harpalus punctatostriatus [1]
Harpalus punctatus [1]
Harpalus pygmaeus [1]
Harpalus retractus [19]
Harpalus reversus [69]
Harpalus rubripes [124]
Harpalus rufipalpis [16]
Harpalus rufipes [105]
Harpalus saxicola [1]
Harpalus schaumii [2]
Harpalus serripes [46]
Harpalus servus [7]
Harpalus signaticornis [21]
Harpalus sinicus [3]
Harpalus smaragdinus [32]
Harpalus solitaris [90]
Harpalus somnulentus [187]
Harpalus sp. [42]
Harpalus sp. 1 [1]
Harpalus sp. 2 [1]
Harpalus sp. DNAS-1E5-197360 [1]
Harpalus sp. DNAS-1E9-197370 [1]
Harpalus sp. NLG-2014 [1]
Harpalus spadiceus [48]
Harpalus stephani [6]
Harpalus subcylindricus [23]
Harpalus sulphuripes [13]
Harpalus tardus [74]
Harpalus tenebrosus [21]
Harpalus texanus [2]
Harpalus tsushimanus [1]
Harpalus vagans [91]
Harpalus ventralis [72]
Harpalus vicarius [1]
Harpalus wagneri [2]
Harpalus xanthopus [5]
Harpalus zabroides [4]
Taxon Description (Wikipedia)
full article at Wikipedia
BOLD Stats
Specimen Records:
4,001
Public Records:
3,342
Specimens with Sequences:
3,541
Public Species:
118
Specimens with Barcodes:
3,252
Public BINs:
92
Species:
122
Species With Barcodes:
102
Species List - Progress
Access Published & Released Data
Contributors (Specimens & Sequencing)
Specimen Depositories:
Sequencing Labs:
Imagery
images representing subtaxa of Harpalus
(
Harpalus zabroides
)
Sample ID:
GBOLRL0454
License:
CreativeCommons - Attribution Non-Commercial Share-Alike (2016)
License Holder:
SNSB, Staatliche Naturwissenschaftliche Sammlungen Bayerns, ZSM (SNSB, Zoologische Staatssammlung Muenchen)
Collection Sites
Collected from 23 countries.
Top 20:
Show All Countries
United States
2285
Finland
19
Bulgaria
3
Germany
626
Pakistan
15
South Korea
3
Canada
326
China
14
India
2
Austria
228
Netherlands
12
Czech Republic
2
France
39
United Kingdom
11
Belgium
2
Norway
35
Hungary
4
Kenya
1
Croatia
27
Italy
3
![(Harpalus D3_D - NEONTcarabid6010) @12 [ ] Copyright (2012) Baldwin, TA National Ecological Observatory Network (NEON) http://www.neoninc.org/content/copyright](http://v3.boldsystems.org/pics/_w300/NEONV/carabid6010e%2B1351525206.jpg "(Harpalus D3_D - NEONTcarabid6010) {} [ ] Copyright (2012)")
![(Harpalus affinis - 10PCCOL-1100) @15 [ ] CreativeCommons - Attribution (2010) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/BBCCM/IMG_3178%2B1291816770.jpg "(Harpalus affinis - 10PCCOL-1100) {} [ ] CreativeCommons - Attribution (2010)")
![(Harpalus albanicus - GBOLRL0457) @11 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2016) SNSB, Staatliche Naturwissenschaftliche Sammlungen Bayerns ZSM (SNSB, Zoologische Staatssammlung Muenchen)](http://v3.boldsystems.org/pics/_w300/GBMUS/GBOLRL0457%2B1502467560.JPG "(Harpalus albanicus - GBOLRL0457) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2016)")
![(Harpalus amputatus - NEON.BET.D10.000377) @14 [ ] Copyright (2010) National Ecological Observatory Network, Inc. National Ecological Observatory Network (NEON) http://www.neoninc.org/content/copyright](http://v3.boldsystems.org/pics/_w300/IUP/Car.D10.000377e%2B1390597876.jpg "(Harpalus amputatus - NEON.BET.D10.000377) {} [ ] Copyright (2010)")
![(Harpalus animosus - NEONTcarabid4261) @13 [ ] Copyright (2010) Blevins, KK and Travers, PD National Ecological Observatory Network (NEON) http://www.neoninc.org/content/copyright](http://v3.boldsystems.org/pics/_w300/NEONU/carabid4261%2B1315515844.jpg "(Harpalus animosus - NEONTcarabid4261) {} [ ] Copyright (2010)")
![(Harpalus anxius - BFB_Col_FK_10685) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015) SNSB, Zoologische Staatssammlung Muenchen SNSB, Zoologische Staatssammlung Muenchen](http://v3.boldsystems.org/pics/_w300/FBCON/BFB_Col_FK_10685%2B1366468924.jpg "(Harpalus anxius - BFB_Col_FK_10685) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015)")
![(Harpalus atratus - BFB_Col_FK_10415) @14 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015) SNSB, Zoologische Staatssammlung Muenchen SNSB, Zoologische Staatssammlung Muenchen](http://v3.boldsystems.org/pics/_w300/FBCOQ/BFB_Col_FK_10415%2B1367955780.jpg "(Harpalus atratus - BFB_Col_FK_10415) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015)")
![(Harpalus attenuatus - BCZSM_COLA_01388) @12 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2012) SNSB, Zoologische Staatssammlung Muenchen SNSB, Zoologische Staatssammlung Muenchen](http://v3.boldsystems.org/pics/_w300/FBCOH/BCZSM_COLA_01388%2B1348244668.JPG "(Harpalus attenuatus - BCZSM_COLA_01388) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2012)")
![(Harpalus autumnalis - GBOL_Col_FK_3326) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015) SNSB, Zoologische Staatssammlung Muenchen SNSB, Zoologische Staatssammlung Muenchen](http://v3.boldsystems.org/pics/_w300/GBCOL/GBOL_Col_FK_3326%2B1350138266.jpg "(Harpalus autumnalis - GBOL_Col_FK_3326) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015)")
![(Harpalus bellieri - BC-LPRCorse1028) @11 [ ] Copyright (2019) Julien Touroult Museum national d'Histoire naturelle, Paris](http://v3.boldsystems.org/pics/_w300/LPRCC/Harpalus_belieri_01%2B1562082014.jpg "(Harpalus bellieri - BC-LPRCorse1028) {} [ ] Copyright (2019)")
![(Harpalus calceatus - BIOUG14399-D05) @15 [ ] CreativeCommons - Attribution (2015) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/GMCHM/BIOUG14399-D05%2B1424280880.jpg "(Harpalus calceatus - BIOUG14399-D05) {} [ ] CreativeCommons - Attribution (2015)")
![(Harpalus caliginosus - NEON.BET.D10.000447) @14 [ ] Copyright (2010) National Ecological Observatory Network, Inc. National Ecological Observatory Network (NEON) http://www.neoninc.org/content/copyright](http://v3.boldsystems.org/pics/_w300/IUP/Car.D10.000447e%2B1390599552.jpg "(Harpalus caliginosus - NEON.BET.D10.000447) {} [ ] Copyright (2010)")
![(Harpalus caspius - GBOL10695) @14 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015) SNSB, Zoologische Staatssammlung Muenchen SNSB, Zoologische Staatssammlung Muenchen](http://v3.boldsystems.org/pics/_w300/GBMIX/GBOL10695%2B1416428998.jpg "(Harpalus caspius - GBOL10695) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015)")
![(Harpalus cautus - CCDB-21414-A07) @11 [ ] CreativeCommons - Attribution (2014) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/LYMAA/CCDB-21414-A07%2B1401366000.jpg "(Harpalus cautus - CCDB-21414-A07) {} [ ] CreativeCommons - Attribution (2014)")
![(Harpalus cephalotes - GBOLRL0518) @11 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2016) SNSB, Staatliche Naturwissenschaftliche Sammlungen Bayerns ZSM (SNSB, Zoologische Staatssammlung Muenchen)](http://v3.boldsystems.org/pics/_w300/GBMUS/GBOLRL0518%2B1502467448.JPG "(Harpalus cephalotes - GBOLRL0518) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2016)")
![(Harpalus compar - CNC COLEO 00118936) @15 [ ] CreativeCommons - Attribution (2011) CNC/CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/CNCCB/CNC_COLEO_00118936%2B1308763610.JPG "(Harpalus compar - CNC COLEO 00118936) {} [ ] CreativeCommons - Attribution (2011)")
![(Harpalus cordifer - CNC COLEO 00118985) @14 [ ] CreativeCommons - Attribution (2011) CNC/CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/CNCCB/CNC_COLEO_00118985%2B1308764272.JPG "(Harpalus cordifer - CNC COLEO 00118985) {} [ ] CreativeCommons - Attribution (2011)")
![(Harpalus cupreus - CAR2140-1) @11 [ ] CC BY SA (2022) Paill Wolfgang Universalmuseum Joanneum, Graz](http://v3.boldsystems.org/pics/_w300/ACAR/CAR2140-1%2B1674161616.JPG "(Harpalus cupreus - CAR2140-1) {} [ ] CC BY SA (2022)")
![(Harpalus desertus - CNC COLEO 00119013) @14 [ ] CreativeCommons - Attribution (2011) CNC/CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/CNCCB/CNC_COLEO_00119013%2B1308765246.JPG "(Harpalus desertus - CNC COLEO 00119013) {} [ ] CreativeCommons - Attribution (2011)")
![(Harpalus dimidiatus - BFB_Col_FK_11410) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015) SNSB, Zoologische Staatssammlung Muenchen SNSB, Zoologische Staatssammlung Muenchen](http://v3.boldsystems.org/pics/_w300/FBCON/BFB_Col_FK_11410%2B1366319818.jpg "(Harpalus dimidiatus - BFB_Col_FK_11410) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015)")
![(Harpalus distinguendus - GBOL 01299) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015) SNSB, Zoologische Staatssammlung Muenchen SNSB, Zoologische Staatssammlung Muenchen](http://v3.boldsystems.org/pics/_w300/GBCOU/GBOL01299%2B1372360156.jpg "(Harpalus distinguendus - GBOL 01299) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015)")
![(Harpalus ellipsis - NEON.BET.D10.000719) @13 [ ] Copyright (2010) National Ecological Observatory Network, Inc. National Ecological Observatory Network (NEON) http://www.neoninc.org/content/copyright](http://v3.boldsystems.org/pics/_w300/IUP/Car.D10.000719e%2B1390855728.jpg "(Harpalus ellipsis - NEON.BET.D10.000719) {} [ ] Copyright (2010)")
![(Harpalus erraticus - CNC COLEO 00118946) @13 [ ] CreativeCommons - Attribution (2011) CNC/CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/CNCCB/CNC_COLEO_00118946%2B1308763694.JPG "(Harpalus erraticus - CNC COLEO 00118946) {} [ ] CreativeCommons - Attribution (2011)")
![(Harpalus erythropus - CNC COLEO 00118939) @14 [ ] CreativeCommons - Attribution (2011) CNC/CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/CNCCB/CNC_COLEO_00118939%2B1308763634.JPG "(Harpalus erythropus - CNC COLEO 00118939) {} [ ] CreativeCommons - Attribution (2011)")
![(Harpalus faunus - CCDB-21414-C01) @15 [ ] CreativeCommons - Attribution (2014) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/LYMAA/CCDB-21414-C01%2B1401366000.jpg "(Harpalus faunus - CCDB-21414-C01) {} [ ] CreativeCommons - Attribution (2014)")
![(Harpalus flavescens - DZMB_Carabidae_0001) @11 [ ] CreativeCommons - Attribution Share-Alike (2016) Michael Raupach DZMB / Senckenberg am Meer](http://v3.boldsystems.org/pics/_w300/EUCAR/DZMB_Carabidae_0001%2B1461605714.jpg "(Harpalus flavescens - DZMB_Carabidae_0001) {} [ ] CreativeCommons - Attribution Share-Alike (2016)")
![(Harpalus flavicornis - GBOLRL0459) @11 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2016) SNSB, Staatliche Naturwissenschaftliche Sammlungen Bayerns ZSM (SNSB, Zoologische Staatssammlung Muenchen)](http://v3.boldsystems.org/pics/_w300/GBMUS/GBOLRL0459%2B1502467566.JPG "(Harpalus flavicornis - GBOLRL0459) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2016)")
![(Harpalus fraternus - NEONTcarabid4259) @14 [ ] Copyright (2010) Blevins, KK and Travers, PD National Ecological Observatory Network (NEON) http://www.neoninc.org/content/copyright](http://v3.boldsystems.org/pics/_w300/NEONU/carabid4259%2B1315516070.jpg "(Harpalus fraternus - NEONTcarabid4259) {} [ ] Copyright (2010)")
![(Harpalus froelichii - BCZSM_COLA_01597) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2012) SNSB, Zoologische Staatssammlung Muenchen SNSB, Zoologische Staatssammlung Muenchen](http://v3.boldsystems.org/pics/_w300/FBCOH/BCZSM_COLA_01597%2B1350412280.JPG "(Harpalus froelichii - BCZSM_COLA_01597) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2012)")
![(Harpalus fulvilabris - CCDB-23051-H08) @15 [ ] CreativeCommons - Attribution (2015) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/COLNF/CCDB-23051-H08%2B1422642652.jpg "(Harpalus fulvilabris - CCDB-23051-H08) {} [ ] CreativeCommons - Attribution (2015)")
![(Harpalus fuscipalpis - 09BBCOL-0673) @14 [ ] CreativeCommons - Attribution (2009) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/USCOL/IMG_8618%2B1250514742.JPG "(Harpalus fuscipalpis - 09BBCOL-0673) {} [ ] CreativeCommons - Attribution (2009)")
![(Harpalus gravis - CCDB-21414-C11) @11 [ ] CreativeCommons - Attribution (2014) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/LYMAA/CCDB-21414-C11%2B1401366000.jpg "(Harpalus gravis - CCDB-21414-C11) {} [ ] CreativeCommons - Attribution (2014)")
![(Harpalus griseus - GBOL_Col_FK_2030) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015) SNSB, Zoologische Staatssammlung Muenchen SNSB, Zoologische Staatssammlung Muenchen](http://v3.boldsystems.org/pics/_w300/GBCOC/GBOL_Col_FK_2030%2B1343422556.jpg "(Harpalus griseus - GBOL_Col_FK_2030) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015)")
![(Harpalus herbivagus - CNC COLEO 00118994) @14 [ ] CreativeCommons - Attribution (2011) CNC/CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/CNCCB/CNC_COLEO_00118994%2B1308764384.JPG "(Harpalus herbivagus - CNC COLEO 00118994) {} [ ] CreativeCommons - Attribution (2011)")
![(Harpalus hirtipes - BCZSM_COLA_01335) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2012) SNSB, Zoologische Staatssammlung Muenchen SNSB, Zoologische Staatssammlung Muenchen](http://v3.boldsystems.org/pics/_w300/FBCOH/BCZSM_COLA_01335%2B1348244668.JPG "(Harpalus hirtipes - BCZSM_COLA_01335) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2012)")
![(Harpalus honestus - BC ZSM COL 00853) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2010) SNSB, Zoologische Staatssammlung Muenchen SNSB, Zoologische Staatssammlung Muenchen](http://v3.boldsystems.org/pics/_w300/FBCOA/BCZSMCOL00853%2B1279639186.JPG "(Harpalus honestus - BC ZSM COL 00853) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2010)")
![(Harpalus indianus - CNC COLEO 00118948) @14 [ ] CreativeCommons - Attribution (2011) CNC/CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/CNCCB/CNC_COLEO_00118948%2B1308763732.JPG "(Harpalus indianus - CNC COLEO 00118948) {} [ ] CreativeCommons - Attribution (2011)")
![(Harpalus indigens - CCDB-21414-D05) @15 [ ] CreativeCommons - Attribution (2014) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/LYMAA/CCDB-21414-D05%2B1401366000.jpg "(Harpalus indigens - CCDB-21414-D05) {} [ ] CreativeCommons - Attribution (2014)")
![(Harpalus innocuus - CNC COLEO 00118999) @12 [ ] CreativeCommons - Attribution (2011) CNC/CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/CNCCB/CNC_COLEO_00118999%2B1308765462.JPG "(Harpalus innocuus - CNC COLEO 00118999) {} [ ] CreativeCommons - Attribution (2011)")
![(Harpalus karamani - BGE_00249_C02) @11 [ ] Creative Commons Attribution Non-Commercial Share-Alike (2024) NTNU University Museum, Department of Natural History NTNU University Museum, Department of Natural History](http://v3.boldsystems.org/pics/_w300/BSNTN/BGE_00249_C02%2B1708464774.jpg "(Harpalus karamani - BGE_00249_C02) {} [ ] Creative Commons Attribution Non-Commercial Share-Alike (2024)")
![(Harpalus katiae - NEONcarabid8572) @15 [ ] Copyright (2010) National Ecological Observatory Network, Inc. National Ecological Observatory Network (NEON) http://www.neoninc.org/content/copyright](http://v3.boldsystems.org/pics/_w300/ISUP/NEON8572E%2B1417811596.jpg "(Harpalus katiae - NEONcarabid8572) {} [ ] Copyright (2010)")
![(Harpalus laevipes - CCDB-23051-G08) @14 [ ] CreativeCommons - Attribution (2015) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/COLNF/CCDB-23051-G08%2B1422642048.jpg "(Harpalus laevipes - CCDB-23051-G08) {} [ ] CreativeCommons - Attribution (2015)")
![(Harpalus laticeps - CNC COLEO 00118965) @15 [ ] CreativeCommons - Attribution (2011) CNC/CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/CNCCB/CNC_COLEO_00118965%2B1308764042.JPG "(Harpalus laticeps - CNC COLEO 00118965) {} [ ] CreativeCommons - Attribution (2011)")
![(Harpalus latus - BCZSM_COL_WL_0081) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2010) Stefan Schmidt SNSB, Zoologische Staatssammlung Muenchen](http://v3.boldsystems.org/pics/_w300/FBCOD/BCZSMCOLWL0081%2B1295265748.JPG "(Harpalus latus - BCZSM_COL_WL_0081) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2010)")
![(Harpalus lewisii - CNC COLEO 00118968) @13 [ ] CreativeCommons - Attribution (2011) CNC/CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/CNCCB/CNC_COLEO_00118968%2B1308764060.JPG "(Harpalus lewisii - CNC COLEO 00118968) {} [ ] CreativeCommons - Attribution (2011)")
![(Harpalus luteicornis - BFB_Col_FK_9875) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015) SNSB, Zoologische Staatssammlung Muenchen SNSB, Zoologische Staatssammlung Muenchen](http://v3.boldsystems.org/pics/_w300/FBCOP/BFB_Col_FK_9875%2B1369150028.jpg "(Harpalus luteicornis - BFB_Col_FK_9875) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015)")
![(Harpalus marginellus - GBOLRL0456) @11 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2016) SNSB, Staatliche Naturwissenschaftliche Sammlungen Bayerns ZSM (SNSB, Zoologische Staatssammlung Muenchen)](http://v3.boldsystems.org/pics/_w300/GBMUS/GBOLRL0456%2B1502467558.JPG "(Harpalus marginellus - GBOLRL0456) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2016)")
![(Harpalus megacephalus - CNC COLEO 00118982) @13 [ ] CreativeCommons - Attribution (2011) CNC/CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/CNCCB/CNC_COLEO_00118982%2B1308056738.JPG "(Harpalus megacephalus - CNC COLEO 00118982) {} [ ] CreativeCommons - Attribution (2011)")
![(Harpalus melancholicus - BC ZSM COL 02140) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2010) Stefan Schmidt SNSB, Zoologische Staatssammlung Muenchen](http://v3.boldsystems.org/pics/_w300/FBCOC/BCZSMCOL02140%2B1286287962.JPG "(Harpalus melancholicus - BC ZSM COL 02140) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2010)")
![(Harpalus modestus - BCZSM_COLA_01577) @11 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2012) SNSB, Zoologische Staatssammlung Muenchen SNSB, Zoologische Staatssammlung Muenchen](http://v3.boldsystems.org/pics/_w300/FBCOH/BCZSM_COLA_01577%2B1350412280.JPG "(Harpalus modestus - BCZSM_COLA_01577) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2012)")
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![(Harpalus nigritarsis - 07PROBE-05025) @14 [ ] CreativeCommons - Attribution (2010) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/MZCHU/IMG_9784%2B1192113952.JPG "(Harpalus nigritarsis - 07PROBE-05025) {} [ ] CreativeCommons - Attribution (2010)")
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![(Harpalus obnixus - CNC COLEO 00119002) @11 [ ] CreativeCommons - Attribution (2011) CNC/CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/CNCCB/CNC_COLEO_00119002%2B1308765422.JPG "(Harpalus obnixus - CNC COLEO 00119002) {} [ ] CreativeCommons - Attribution (2011)")
![(Harpalus ochropus - CBG-A08781-A11) @11 [ ] CreativeCommons - Attribution Share - Alike (2023) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/YBIVV/CBG-A08781-A11%2B1692628272.jpg "(Harpalus ochropus - CBG-A08781-A11) {} [ ] CreativeCommons - Attribution Share - Alike (2023)")
![(Harpalus opacipennis - 09BBCOL-0672) @16 [ ] CreativeCommons - Attribution (2009) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/USCOL/IMG_8617%2B1250514732.JPG "(Harpalus opacipennis - 09BBCOL-0672) {} [ ] CreativeCommons - Attribution (2009)")
![(Harpalus paratus - NEON.BET.D10.000669) @13 [ ] Copyright (2010) National Ecological Observatory Network, Inc. National Ecological Observatory Network (NEON) http://www.neoninc.org/content/copyright](http://v3.boldsystems.org/pics/_w300/IUP/Car.D10.000669e%2B1390852886.jpg "(Harpalus paratus - NEON.BET.D10.000669) {} [ ] Copyright (2010)")
![(Harpalus pensylvanicus - BIOUG02010-F11) @15 [ ] CreativeCommons - Attribution (2012) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/BBCCA/IMG_8817%2B1332939728.JPG "(Harpalus pensylvanicus - BIOUG02010-F11) {} [ ] CreativeCommons - Attribution (2012)")
![(Harpalus picipennis - BCZSM_COLA_01631) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015) SNSB, Zoologische Staatssammlung Muenchen SNSB, Zoologische Staatssammlung Muenchen](http://v3.boldsystems.org/pics/_w300/FBCOJ/BCZSM_COLA_01631%2B1355953990.jpg "(Harpalus picipennis - BCZSM_COLA_01631) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015)")
![(Harpalus plenalis - CNC COLEO 00119008) @14 [ ] CreativeCommons - Attribution (2011) CNC/CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/CNCCB/CNC_COLEO_00119008%2B1308765324.JPG "(Harpalus plenalis - CNC COLEO 00119008) {} [ ] CreativeCommons - Attribution (2011)")
![(Harpalus politus - 39_Fa2_be001) @11 [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2023) Forest Ecology Course TU Darmstadt](http://v3.boldsystems.org/pics/_w300/WALDD/Pl39_Fa2_Be001_dorsal%2B1688742922.jpg "(Harpalus politus - 39_Fa2_be001) {} [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2023)")
![(Harpalus progrediens - GBOLRL0521) @11 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2016) SNSB, Staatliche Naturwissenschaftliche Sammlungen Bayerns ZSM (SNSB, Zoologische Staatssammlung Muenchen)](http://v3.boldsystems.org/pics/_w300/GBMUS/GBOLRL0521%2B1502467454.JPG "(Harpalus progrediens - GBOLRL0521) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2016)")
![(Harpalus protractus - BIOUG02010-F01) @14 [ ] CreativeCommons - Attribution (2012) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/BBCCA/IMG_8807%2B1332771594.JPG "(Harpalus protractus - BIOUG02010-F01) {} [ ] CreativeCommons - Attribution (2012)")
![(Harpalus providens - CCDB-21414-F07) @14 [ ] CreativeCommons - Attribution (2014) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/LYMAA/CCDB-21414-F07%2B1401366060.JPG "(Harpalus providens - CCDB-21414-F07) {} [ ] CreativeCommons - Attribution (2014)")
![(Harpalus pumilus - GBOL_Col_FK_7878) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015) SNSB, Zoologische Staatssammlung Muenchen SNSB, Zoologische Staatssammlung Muenchen](http://v3.boldsystems.org/pics/_w300/GBCOU/GBOL_Col_FK_7878%2B1379354984.jpg "(Harpalus pumilus - GBOL_Col_FK_7878) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015)")
![(Harpalus punctatus - BCZSM_COLA_01357) @11 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2012) SNSB, Zoologische Staatssammlung Muenchen SNSB, Zoologische Staatssammlung Muenchen](http://v3.boldsystems.org/pics/_w300/FBCOH/BCZSM_COLA_01357%2B1348244668.JPG "(Harpalus punctatus - BCZSM_COLA_01357) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2012)")
![(Harpalus pygmaeus - BGE_00249_C04) @11 [ ] Creative Commons Attribution Non-Commercial Share-Alike (2024) NTNU University Museum, Department of Natural History NTNU University Museum, Department of Natural History](http://v3.boldsystems.org/pics/_w300/BSNTN/BGE_00249_C04%2B1708464892.jpg "(Harpalus pygmaeus - BGE_00249_C04) {} [ ] Creative Commons Attribution Non-Commercial Share-Alike (2024)")
![(Harpalus retractus - NEONTcarabid5139) @11 [ ] Copyright (2012) Blevins, KK and Travers, PD National Ecological Observatory Network (NEON) http://www.neoninc.org/content/copyright](http://v3.boldsystems.org/pics/_w300/NEONU/carabid5139%2B1324499328.JPG "(Harpalus retractus - NEONTcarabid5139) {} [ ] Copyright (2012)")
![(Harpalus reversus - NEONTcarabid690) @13 [ ] Copyright (2010) Blevins, KK and Travers, PD National Ecological Observatory Network (NEON) http://www.neoninc.org/content/copyright](http://v3.boldsystems.org/pics/_w300/NEONT/carabid690%2B1284027424.jpg "(Harpalus reversus - NEONTcarabid690) {} [ ] Copyright (2010)")
![(Harpalus rubripes - ZMUO.005711) @14 [ ] CreativeCommons - Attribution Non-Commercial (2012) Marko Mutanen University of Oulu](http://v3.boldsystems.org/pics/_w300/COLFF/ZMUO.005711%2B1365635356.JPG "(Harpalus rubripes - ZMUO.005711) {} [ ] CreativeCommons - Attribution Non-Commercial (2012)")
![(Harpalus rufipalpis - BFB_Col_FK_11045) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015) SNSB, Zoologische Staatssammlung Muenchen SNSB, Zoologische Staatssammlung Muenchen](http://v3.boldsystems.org/pics/_w300/FBCOP/BFB_Col_FK_11045%2B1366817322.jpg "(Harpalus rufipalpis - BFB_Col_FK_11045) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015)")
![(Harpalus rufipes - ZMUO.001005) @15 [ ] CreativeCommons - Attribution Non-Commercial (2012) Marko Mutanen University of Oulu](http://v3.boldsystems.org/pics/_w300/COLFB/ZMUO.001005%2B1329405646.JPG "(Harpalus rufipes - ZMUO.001005) {} [ ] CreativeCommons - Attribution Non-Commercial (2012)")
![(Harpalus serripes - GBOL_Col_FK_7400) @14 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015) SNSB, Zoologische Staatssammlung Muenchen SNSB, Zoologische Staatssammlung Muenchen](http://v3.boldsystems.org/pics/_w300/GBCOU/GBOL_Col_FK_7400%2B1378415416.jpg "(Harpalus serripes - GBOL_Col_FK_7400) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015)")
![(Harpalus servus - BCZSM_COLA_00841) @12 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2010) SNSB, Zoologische Staatssammlung Muenchen SNSB, Zoologische Staatssammlung Muenchen](http://v3.boldsystems.org/pics/_w300/FBCOD/BC_ZSM_COLA_00841%2B1312292676.jpg "(Harpalus servus - BCZSM_COLA_00841) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2010)")
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![(Harpalus solitaris - ZMUO.004230) @13 [ ] CreativeCommons - Attribution Non-Commercial (2012) Marko Mutanen University of Oulu](http://v3.boldsystems.org/pics/_w300/COLFD/ZMUO.004230%2B1347030422.JPG "(Harpalus solitaris - ZMUO.004230) {} [ ] CreativeCommons - Attribution Non-Commercial (2012)")
![(Harpalus somnulentus - CCDB-23051-G10) @15 [ ] CreativeCommons - Attribution (2015) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/COLNF/CCDB-23051-G10%2B1422642068.jpg "(Harpalus somnulentus - CCDB-23051-G10) {} [ ] CreativeCommons - Attribution (2015)")
![(Harpalus sp - NEON.BET.D03.000215) @12 [ ] Copyright (2010) National Ecological Observatory Network, Inc. National Ecological Observatory Network (NEON) http://www.neoninc.org/content/copyright](http://v3.boldsystems.org/pics/_w300/IUP/BET.D03.000215e%2B1395767786.jpg "(Harpalus sp - NEON.BET.D03.000215) {} [ ] Copyright (2010)")
![(Harpalus sp. 1 - NEONTcarabid568) @11 [ ] Copyright (2010) Blevins, KK and Travers, PD National Ecological Observatory Network (NEON) http://www.neoninc.org/content/copyright](http://v3.boldsystems.org/pics/_w300/NEONT/carabid568%2B1283972794.jpg "(Harpalus sp. 1 - NEONTcarabid568) {} [ ] Copyright (2010)")
![(Harpalus sp. 2 - NEONTcarabid1495) @13 [ ] Copyright (2010) Blevins, KK and Travers, PD National Ecological Observatory Network (NEON) http://www.neoninc.org/content/copyright](http://v3.boldsystems.org/pics/_w300/NEONT/carabid1495%2B1283971770.jpg "(Harpalus sp. 2 - NEONTcarabid1495) {} [ ] Copyright (2010)")
![(Harpalus spadiceus - NEONcarabid8253) @13 [ ] Copyright (2010) National Ecological Observatory Network, Inc. National Ecological Observatory Network (NEON) http://www.neoninc.org/content/copyright](http://v3.boldsystems.org/pics/_w300/ISUP/NEON8253E%2B1417554466.jpg "(Harpalus spadiceus - NEONcarabid8253) {} [ ] Copyright (2010)")
![(Harpalus subcylindricus - BFB_Col_FK_9650) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015) SNSB, Zoologische Staatssammlung Muenchen SNSB, Zoologische Staatssammlung Muenchen](http://v3.boldsystems.org/pics/_w300/FBCOK/BFB_Col_FK_9650%2B1357160740.jpg "(Harpalus subcylindricus - BFB_Col_FK_9650) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015)")
![(Harpalus sulphuripes - LPRC2021-0997) @11 [ ] By-SA Creative Common (2021) Rodolphe Rougerie Museum national d'Histoire naturelle, Paris](http://v3.boldsystems.org/pics/_w300/LPRCI/BJZ_1062%2B1623513328.JPG "(Harpalus sulphuripes - LPRC2021-0997) {} [ ] By-SA Creative Common (2021)")
![(Harpalus tardus - ZMUO.001617) @14 [ ] CreativeCommons - Attribution Non-Commercial (2012) Marko Mutanen University of Oulu](http://v3.boldsystems.org/pics/_w300/COLFB/ZMUO.001617%2B1331912976.JPG "(Harpalus tardus - ZMUO.001617) {} [ ] CreativeCommons - Attribution Non-Commercial (2012)")
![(Harpalus tenebrosus - BFB_Col_FK_10260) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015) SNSB, Zoologische Staatssammlung Muenchen SNSB, Zoologische Staatssammlung Muenchen](http://v3.boldsystems.org/pics/_w300/FBCOP/BFB_Col_FK_10260%2B1369086348.jpg "(Harpalus tenebrosus - BFB_Col_FK_10260) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015)")
![(Harpalus vagans - CNC COLEO 00118935) @13 [ ] CreativeCommons - Attribution (2011) CNC/CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/CNCCB/CNC_COLEO_00118935%2B1308763600.JPG "(Harpalus vagans - CNC COLEO 00118935) {} [ ] CreativeCommons - Attribution (2011)")
![(Harpalus ventralis - 09BBCOL-0667) @14 [ ] CreativeCommons - Attribution (2009) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/USCOL/IMG_4985%2B1250281622.JPG "(Harpalus ventralis - 09BBCOL-0667) {} [ ] CreativeCommons - Attribution (2009)")
![(Harpalus xanthopus - ZMUO.004899) @14 [ ] CreativeCommons - Attribution Non-Commercial (2012) Marko Mutanen University of Oulu](http://v3.boldsystems.org/pics/_w300/COLFE/ZMUO.004899%2B1358795816.JPG "(Harpalus xanthopus - ZMUO.004899) {} [ ] CreativeCommons - Attribution Non-Commercial (2012)")
![(Harpalus zabroides - GBOLRL0454) @11 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2016) SNSB, Staatliche Naturwissenschaftliche Sammlungen Bayerns ZSM (SNSB, Zoologische Staatssammlung Muenchen)](http://v3.boldsystems.org/pics/_w300/GBMUS/GBOLRL0454%2B1502467552.JPG "(Harpalus zabroides - GBOLRL0454) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2016)")
