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Latest Version
Harpalinae {subfamily} -
Arthropoda
;
Insecta
;
Coleoptera
;
Carabidae
;
Print
For the latest taxonomy data, visit
boldsystems.org
.
Search Taxonomy
Sub-taxa
Tribes (37)
Abacetini [76]
Anthiini [20]
Atranini [8]
Calophaenini [9]
Chlaeniini [1307]
Cnemalobini [22]
Collyridini [31]
Corsyrini [8]
Ctenodactylini [22]
Cyclosomini [227]
Dercylini [29]
Drimostomatini [2]
Dryptini [30]
Galeritini [311]
Harpalini [11092]
Helluonini [235]
Hexagoniini [4]
Lachnophorini [56]
Lebiini [3756]
Licinini [1363]
Masoreini [27]
Metiini [52]
Morionini [23]
Odacanthini [117]
Oodini [208]
Orthogoniini [8]
Panagaeini [110]
Peleciini [7]
Pentagonicini [262]
Perigonini [65]
Physocrotaphini [2]
Platynini [4662]
Pseudomorphini [27]
Pterostichini [10573]
Sphodrini [2693]
Zabrini [4480]
Zuphiini [83]
Genera (1)
Selenophori [1]
Taxon Description (Wikipedia)
full article at Wikipedia
BOLD Stats
Specimen Records:
44,951
Public Records:
35,217
Specimens with Sequences:
39,677
Public Species:
1,872
Specimens with Barcodes:
36,244
Public BINs:
1,636
Species:
2,159
Species With Barcodes:
1,604
Species List - Progress
Access Published & Released Data
Contributors (Specimens & Sequencing)
Specimen Depositories:
Sequencing Labs:
Imagery
images representing subtaxa of Harpalinae
(
Zuphiini
)
Sample ID:
CNC COLEO 00119546
License:
CreativeCommons - Attribution (2011)
License Holder:
CNC/CBG Photography Group, Centre for Biodiversity Genomics
Collection Sites
Collected from 77 countries.
Top 20:
Show All Countries
United States
19980
Norway
265
Colombia
135
Canada
5537
United Kingdom
222
Ecuador
125
Germany
4298
Netherlands
208
Croatia
119
Costa Rica
2421
Australia
183
Finland
107
Austria
2099
Peru
165
South Africa
107
Ghana
515
Pakistan
161
Greece
65
France
350
Bulgaria
153
![(Abacetini - CCDB-21417-D05) @14 [ ] CreativeCommons - Attribution (2014) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/LYMAA/ccdb-21417-d05%2B1401386178.jpg "(Abacetini - CCDB-21417-D05) {} [ ] CreativeCommons - Attribution (2014)")
![(Anthiini - NIBGE COL-01379) @14 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2011) Muhammad Ashfaq National Institute for Biotechnology and Genetic Engineering Faisalabad Pakistan](http://v3.boldsystems.org/pics/_w300/MACOL/NIBGE_COL-01379%2B1311036506.JPG "(Anthiini - NIBGE COL-01379) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2011)")
![(Atranini - CCDB-21331-E10) @15 [ ] CreativeCommons - Attribution (2014) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/LYMAA/ccdb-21331-e10%2B1397228426.jpg "(Atranini - CCDB-21331-E10) {} [ ] CreativeCommons - Attribution (2014)")
![(Calophaenini - BC-GF2024-0711) @11 [ ] Creative Commons (2024) rodolphe rougerie Museum national d'histoire naturelle](http://v3.boldsystems.org/pics/_w300/FGBEL/GF_0711%2B315525600.JPG "(Calophaenini - BC-GF2024-0711) {} [ ] Creative Commons (2024)")
![(Chlaeniini - BCZSM_COLA_01547) @16 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2012) SNSB, Zoologische Staatssammlung Muenchen SNSB, Zoologische Staatssammlung Muenchen](http://v3.boldsystems.org/pics/_w300/FBCOH/BCZSM_COLA_01547%2B1350412280.JPG "(Chlaeniini - BCZSM_COLA_01547) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2012)")
![(Cnemalobini - CCDB-30443-A02) @11 [ ] CreativeCommons - Attribution (2018) Smithsonian Institution Smithsonian Institution](http://v3.boldsystems.org/pics/_w300/SICOA/CCDB-30443-A02_Dorsal%2B1540910400.jpg "(Cnemalobini - CCDB-30443-A02) {} [ ] CreativeCommons - Attribution (2018)")
![(Collyridini - NIBGE COL-01149) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2011) Muhammad Ashfaq National Institute for Biotechnology and Genetic Engineering Faisalabad Pakistan](http://v3.boldsystems.org/pics/_w300/MACOL/NIBGE_COL-1149%2B1307714192.jpg "(Collyridini - NIBGE COL-01149) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2011)")
![(Corsyrini - CCDB-30443-C04) @11 [ ] CreativeCommons - Attribution (2018) Smithsonian Institution Smithsonian Institution](http://v3.boldsystems.org/pics/_w300/SICOA/CCDB-30443-C04_Dorsal%2B1542137280.jpg "(Corsyrini - CCDB-30443-C04) {} [ ] CreativeCommons - Attribution (2018)")
![(Ctenodactylini - 09BBCOL-0958) @14 [ ] CreativeCommons - Attribution (2009) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/USCOL/IMG_9291%2B1250624936.JPG "(Ctenodactylini - 09BBCOL-0958) {} [ ] CreativeCommons - Attribution (2009)")
![(Cyclosomini - BIOUG02279-D11) @14 [ ] CreativeCommons - Attribution (2012) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/BBCCA/bioug02279-d11%2B1354034264.JPG "(Cyclosomini - BIOUG02279-D11) {} [ ] CreativeCommons - Attribution (2012)")
![(Dercylini - CCDB-30447-A09) @11 [ ] CreativeCommons - Attribution (2018) Smithsonian Institution Smithsonian Institution](http://v3.boldsystems.org/pics/_w300/SICOA/CCDB-30447-A09_Lateral%2B1536860040.jpg "(Dercylini - CCDB-30447-A09) {} [ ] CreativeCommons - Attribution (2018)")
![(Drimostomatini - CCDB-30450-G08) @11 [ ] CreativeCommons - Attribution (2018) Smithsonian Institution Smithsonian Institution](http://v3.boldsystems.org/pics/_w300/SICOA/CCDB-30450-G08_Dorsal%2B1539098494.jpg "(Drimostomatini - CCDB-30450-G08) {} [ ] CreativeCommons - Attribution (2018)")
![(Dryptini - BC ZSM COL 01911) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2010) Stefan Schmidt SNSB, Zoologische Staatssammlung Muenchen](http://v3.boldsystems.org/pics/_w300/FBCOC/BCZSMCOL01911%2B1284714370.JPG "(Dryptini - BC ZSM COL 01911) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2010)")
![(Galeritini - CCDB-21430-D07) @15 [ ] CreativeCommons - Attribution (2014) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/LYMAA/CCDB-21430-D07%2B1406035248.JPG "(Galeritini - CCDB-21430-D07) {} [ ] CreativeCommons - Attribution (2014)")
![(Harpalini - 09BBCOL-0669) @16 [ ] CreativeCommons - Attribution (2009) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/USCOL/IMG_8614%2B1250514704.JPG "(Harpalini - 09BBCOL-0669) {} [ ] CreativeCommons - Attribution (2009)")
![(Helluonini - 10BBCOL-0116) @15 [ ] CreativeCommons - Attribution (2010) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/USCOC/10bbcol-0116%2B1279553772.jpg "(Helluonini - 10BBCOL-0116) {} [ ] CreativeCommons - Attribution (2010)")
![(Hexagoniini - CCDB-30450-A05) @11 [ ] CreativeCommons - Attribution (2018) Smithsonian Institution Smithsonian Institution](http://v3.boldsystems.org/pics/_w300/SICOA/CCDB-30450-A05_Dorsal%2B1537972320.jpg "(Hexagoniini - CCDB-30450-A05) {} [ ] CreativeCommons - Attribution (2018)")
![(Lachnophorini - NEONTcarabid5581) @15 [ ] Copyright (2012) Barton, M National Ecological Observatory Network (NEON) http://www.neoninc.org/content/copyright](http://v3.boldsystems.org/pics/_w300/NEONT/carabid5581%2B1369403022.jpg "(Lachnophorini - NEONTcarabid5581) {} [ ] Copyright (2012)")
![(Lebiini - BIOUG00972-B09) @16 [ ] CreativeCommons - Attribution (2011) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/JSCOL/bioug00972-b09%2B1311770604.jpg "(Lebiini - BIOUG00972-B09) {} [ ] CreativeCommons - Attribution (2011)")
![(Licinini - CCDB-21431-C12) @15 [ ] CreativeCommons - Attribution (2014) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/LYMAA/ccdb-21431-c12%2B1401471714.jpg "(Licinini - CCDB-21431-C12) {} [ ] CreativeCommons - Attribution (2014)")
![(Masoreini - ZFMK_COL_2010_659) @13 [ ] CreativeCommons - Attribution (2011) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/EUCAR/zfmk_col_2010_659%2B1311864908.jpg "(Masoreini - ZFMK_COL_2010_659) {} [ ] CreativeCommons - Attribution (2011)")
![(Metiini - CCDB-30450-B12) @11 [ ] CreativeCommons - Attribution (2018) Smithsonian Institution Smithsonian Institution](http://v3.boldsystems.org/pics/_w300/SICOA/CCDB-30450-B12_Dorsal%2B1537988400.jpg "(Metiini - CCDB-30450-B12) {} [ ] CreativeCommons - Attribution (2018)")
![(Morionini - CCDB-21406-F03) @14 [ ] CreativeCommons - Attribution (2014) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/LYMAA/ccdb-21406-f03%2B1397593060.jpg "(Morionini - CCDB-21406-F03) {} [ ] CreativeCommons - Attribution (2014)")
![(Odacanthini - ZFMK_COL_2011_227) @14 [ ] CreativeCommons - Attribution (2011) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/EUCAR/ccdb-12751-f03%2B1311283620.jpg "(Odacanthini - ZFMK_COL_2011_227) {} [ ] CreativeCommons - Attribution (2011)")
![(Oodini - CCDB-21431-H02) @15 [ ] CreativeCommons - Attribution (2014) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/LYMAA/ccdb-21431-h02%2B1401713120.jpg "(Oodini - CCDB-21431-H02) {} [ ] CreativeCommons - Attribution (2014)")
![(Orthogoniini - CCDB-30450-D09) @11 [ ] CreativeCommons - Attribution (2018) Smithsonian Institution Smithsonian Institution](http://v3.boldsystems.org/pics/_w300/SICOA/CCDB-30450-D09_Lateral%2B1538060220.jpg "(Orthogoniini - CCDB-30450-D09) {} [ ] CreativeCommons - Attribution (2018)")
![(Panagaeini - ZMUO.029353) @15 [ ] CreativeCommons - Attribution Non-Commercial (2017) Marko Mutanen University of Oulu](http://v3.boldsystems.org/pics/_w300/COLFH/ZMUO.029353%2B1496100632.JPG "(Panagaeini - ZMUO.029353) {} [ ] CreativeCommons - Attribution Non-Commercial (2017)")
![(Peleciini - CCDB-30450-D12) @11 [ ] CreativeCommons - Attribution (2018) Smithsonian Institution Smithsonian Institution](http://v3.boldsystems.org/pics/_w300/SICOA/CCDB-30450-D12_Dorsal%2B1538061180.jpg "(Peleciini - CCDB-30450-D12) {} [ ] CreativeCommons - Attribution (2018)")
![(Pentagonicini - BIOUG22791-G09) @14 [ ] CreativeCommons - Attribution (2015) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/GMACF/BIOUG22791-G09%2B1442345580.jpg "(Pentagonicini - BIOUG22791-G09) {} [ ] CreativeCommons - Attribution (2015)")
![(Perigonini - RMNH.INS.535232) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2013) Unspecified Naturalis Biodiversity Center](http://v3.boldsystems.org/pics/_w300/NLCOA/RMNH.INS.535232_dor%2B1445540330.jpg "(Perigonini - RMNH.INS.535232) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2013)")
![(Physocrotaphini - CCDB-30450-E05) @11 [ ] CreativeCommons - Attribution (2018) Smithsonian Institution Smithsonian Institution](http://v3.boldsystems.org/pics/_w300/SICOA/CCDB-30450-E05_Dorsal%2B1538068800.jpg "(Physocrotaphini - CCDB-30450-E05) {} [ ] CreativeCommons - Attribution (2018)")
![(Platynini - CCDB-23523-F10) @15 [ ] CreativeCommons - Attribution (2015) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/LYMAB/CCDB-23523-F10%2B1444754314.jpg "(Platynini - CCDB-23523-F10) {} [ ] CreativeCommons - Attribution (2015)")
![(Pseudomorphini - CCDB-30448-A04) @11 [ ] CreativeCommons - Attribution (2018) Smithsonian Institution Smithsonian Institution](http://v3.boldsystems.org/pics/_w300/SICOA/CCDB-30448-A04_Dorsal%2B1537790340.jpg "(Pseudomorphini - CCDB-30448-A04) {} [ ] CreativeCommons - Attribution (2018)")
![(Pterostichini - CCDB-21431-C04) @15 [ ] CreativeCommons - Attribution (2014) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/LYMAA/ccdb-21431-c04%2B1401471530.jpg "(Pterostichini - CCDB-21431-C04) {} [ ] CreativeCommons - Attribution (2014)")
![(Sphodrini - CNC COLEO 00118598) @16 [ ] CreativeCommons - Attribution (2011) CNC/CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/CNCCB/CNC_COLEO_00118598%2B1307982920.JPG "(Sphodrini - CNC COLEO 00118598) {} [ ] CreativeCommons - Attribution (2011)")
![(Zabrini - BIOUG14142-E05) @15 [ ] CreativeCommons - Attribution (2015) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/SSROB/BIOUG14142-E05%2B1422649230.jpg "(Zabrini - BIOUG14142-E05) {} [ ] CreativeCommons - Attribution (2015)")
![(Zuphiini - CNC COLEO 00119546) @13 [ ] CreativeCommons - Attribution (2011) CNC/CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/CNCCB/CNC_COLEO_00119546%2B1308231618.JPG "(Zuphiini - CNC COLEO 00119546) {} [ ] CreativeCommons - Attribution (2011)")
