Please note: Due to scheduled electrical maintenance by the University of Guelph, BOLD Systems will have reduced analysis capacity from June 6, 2025 12:00 pm until June 9, 2025 10:30 am EDT.
You've been logged out. Click here to log back in.
full article at Wikipedia
![(Smittia alpilonga - PP0290) @11 [ ] CreativeCommons - Attribution Non-Commercial (2019) Jaakko Pohjoismaeki University of Eastern Finland](http://v3.boldsystems.org/pics/_w300/FILOU/PP0290%2B1554995924.jpg "(Smittia alpilonga - PP0290) {} [ ] CreativeCommons - Attribution Non-Commercial (2019)")
![(Smittia aterrima - BIOUG17282-H01) @14 [ ] CreativeCommons - Attribution (2015) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/CNVUB/BIOUG17282-H01%2B1449527214.jpg "(Smittia aterrima - BIOUG17282-H01) {} [ ] CreativeCommons - Attribution (2015)")
![(Smittia betuletorum - ZMUO.025823) @11 [ ] CreativeCommons - Attribution Non-Commercial (2016) Marko Mutanen University of Oulu](http://v3.boldsystems.org/pics/_w300/CHIFI/ZMUO.025823%2B1467750262.JPG "(Smittia betuletorum - ZMUO.025823) {} [ ] CreativeCommons - Attribution Non-Commercial (2016)")
![(Smittia brevifurcata - BGE_00003_F01) @11 [ ] CreativeCommons Attribution Share-Alike (by-sa) (2024) Trustees of the Natural History Museum, London Natural History Museum, London](http://v3.boldsystems.org/pics/_w300/BSNHM/013442720_Smittia_brevifurcata_pinned_specimen%2B1738598162.jpg "(Smittia brevifurcata - BGE_00003_F01) {} [ ] CreativeCommons Attribution Share-Alike (by-sa) (2024)")
![(Smittia brevipennis - SV514) @12 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2011) NTNU Museum of Natural History and Archaeology NTNU Museum of Natural History and Archaeology](http://v3.boldsystems.org/pics/_w300/CHRFI/SV514%2B1304437434.jpg "(Smittia brevipennis - SV514) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2011)")
![(Smittia cf. extrema - BIOUG07863-G11) @14 [ ] CreativeCommons - Attribution (2015) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/GMGLG/BIOUG07863-G11%2B1421275320.jpg "(Smittia cf. extrema - BIOUG07863-G11) {} [ ] CreativeCommons - Attribution (2015)")
![(Smittia cf. stercoraria - TRD-CH121) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2014) NTNU University Museum, Department of Natural History NTNU University Museum, Department of Natural History](http://v3.boldsystems.org/pics/_w300/CHMNO/TRD-CH121%2B1404839208.jpg "(Smittia cf. stercoraria - TRD-CH121) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2014)")
![(Smittia contingens - NHMUK014451689) @11 [ ] CreativeCommons Attribution (by) (2024) The Trustees of the Natural History Museum, London Natural History Museum, London](http://v3.boldsystems.org/pics/_w300/UKAN/014451689_DorsoLateral%2B1709580774.jpg "(Smittia contingens - NHMUK014451689) {} [ ] CreativeCommons Attribution (by) (2024)")
![(Smittia edwardsi - BIOUG08660-C05) @15 [ ] CreativeCommons - Attribution (2014) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/SMTPD/BIOUG08660-C05%2B1415391500.jpg "(Smittia edwardsi - BIOUG08660-C05) {} [ ] CreativeCommons - Attribution (2014)")
![(Smittia extrema - BIOUG07711-D04) @13 [ ] CreativeCommons - Attribution (2015) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/GMGLA/BIOUG07711-D04%2B1427312358.jpg "(Smittia extrema - BIOUG07711-D04) {} [ ] CreativeCommons - Attribution (2015)")
![(Smittia gr._aquatilis - RMNH.5091578) @11 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2018) Unspecified Naturalis Biodiversity Center](http://v3.boldsystems.org/pics/_w300/NLCHR/RMNH.5091578_1%2B1530847228.jpg "(Smittia gr._aquatilis - RMNH.5091578) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2018)")
![(Smittia leucopogon - BIOUG02204-H08) @12 [ ] CreativeCommons - Attribution (2012) Muhammad Ashfaq Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/MAMTD/BIOUG02204-H08%2B1348844950.jpg "(Smittia leucopogon - BIOUG02204-H08) {} [ ] CreativeCommons - Attribution (2012)")
![(Smittia longicosta - SV 730) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2013) NTNU Museum of Natural History and Archaeology NTNU Museum of Natural History and Archaeology](http://v3.boldsystems.org/pics/_w300/CHRSV/SV730%2B1376061252.jpg "(Smittia longicosta - SV 730) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2013)")
![(Smittia nigrata - BGE_00003_D01) @11 [ ] CreativeCommons Attribution Share-Alike (by-sa) (2024) Trustees of the Natural History Museum, London Natural History Museum, London](http://v3.boldsystems.org/pics/_w300/BSNHM/013442723_Smittia_nigrata_pinned_specimen%2B1738598166.jpg "(Smittia nigrata - BGE_00003_D01) {} [ ] CreativeCommons Attribution Share-Alike (by-sa) (2024)")
![(Smittia nudipennis - SOE243) @12 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2010) NTNU Museum of Natural History and Archaeology NTNU Museum of Natural History and Archaeology](http://v3.boldsystems.org/pics/_w300/MIDGE/SOE243%2B1204138448.jpg "(Smittia nudipennis - SOE243) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2010)")
![(Smittia paranudipennis - TRD-CH14) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2013) NTNU Museum of Natural History and Archaeology NTNU Museum of Natural History and Archaeology](http://v3.boldsystems.org/pics/_w300/CHMNO/TRD-CH14%2B1383839060.jpg "(Smittia paranudipennis - TRD-CH14) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2013)")
![(Smittia pratorum - NIESD0175) @12 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015) Chironomid Group, NIES National Institute for Environmental Studies, Japan](http://v3.boldsystems.org/pics/_w300/JCDB/D0175%5B1%5D%2B1448472274.jpg "(Smittia pratorum - NIESD0175) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015)")
![(Smittia reissi - ZSM34343-G07) @12 [ ] Unspecified (default): All Rights Reserved SNSB, Staatliche Naturwissenschaftliche Sammlungen Bayerns ZSM (SNSB, Zoologische Staatssammlung Muenchen)](http://v3.boldsystems.org/pics/_w300/FBCHI/ZSM34343-G07%2B1475240172.jpg "(Smittia reissi - ZSM34343-G07) {} [ ] Unspecified (default): All Rights Reserved")
![(Smittia scutellosetosa - TRD-CH90) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2014) NTNU University Museum, Department of Natural History NTNU University Museum, Department of Natural History](http://v3.boldsystems.org/pics/_w300/CHMNO/TRD-CH90%2B1405002620.jpg "(Smittia scutellosetosa - TRD-CH90) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2014)")
![(Smittia sp - CBG-A18172-E02) @11 [ ] CreativeCommons - Attribution Share - Alike (2023) Centre for Biodiversity Genomics Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/GMLCL/CBG-A18172-E02%2B1697481790.jpg "(Smittia sp - CBG-A18172-E02) {} [ ] CreativeCommons - Attribution Share - Alike (2023)")
![(Smittia sp. 1 SW - BIOUG06062-D06) @12 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015) SNSB, Zoologische Staatssammlung Muenchen SNSB, Zoologische Staatssammlung Muenchen](http://v3.boldsystems.org/pics/_w300/GMGRD/BIOUG06062-D06%2B1418047296.jpg "(Smittia sp. 1 SW - BIOUG06062-D06) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015)")
![(Smittia sp. 10ES - Finnmark162) @14 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2012) NTNU Museum of Natural History and Archaeology NTNU Museum of Natural History and Archaeology](http://v3.boldsystems.org/pics/_w300/CHRFI/Fi162%2B1346868632.jpg "(Smittia sp. 10ES - Finnmark162) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2012)")
![(Smittia sp. 13ES - ATNA479) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2013) NTNU Museum of Natural History and Archaeology NTNU Museum of Natural History and Archaeology](http://v3.boldsystems.org/pics/_w300/ATNA/AT479%2B1378486688.jpg "(Smittia sp. 13ES - ATNA479) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2013)")
![(Smittia sp. 14ES - BIOUG21770-A06) @13 [ ] CreativeCommons - Attribution (2015) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/RRMFB/BIOUG21770-A06%2B1437510320.jpg "(Smittia sp. 14ES - BIOUG21770-A06) {} [ ] CreativeCommons - Attribution (2015)")
![(Smittia sp. 15ES - TRD-CH200) @14 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015) NTNU University Museum, Department of Natural History NTNU University Museum, Department of Natural History](http://v3.boldsystems.org/pics/_w300/CHMNO/TRD-CH200%2B1427207980.jpg "(Smittia sp. 15ES - TRD-CH200) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015)")
![(Smittia sp. 16ES - 08WOLVES-01242) @15 [ ] CreativeCommons - Attribution (2012) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/DARC/bioug00539-a07%2B1328192832.JPG "(Smittia sp. 16ES - 08WOLVES-01242) {} [ ] CreativeCommons - Attribution (2012)")
![(Smittia sp. 17ES - 08WOLVES-01271) @13 [ ] CreativeCommons - Attribution (2012) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/DARC/bioug00539-c12%2B1328194906.JPG "(Smittia sp. 17ES - 08WOLVES-01271) {} [ ] CreativeCommons - Attribution (2012)")
![(Smittia sp. 18ES - Finnmark550) @12 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2012) NTNU Museum of Natural History and Archaeology NTNU Museum of Natural History and Archaeology](http://v3.boldsystems.org/pics/_w300/CHRFI/FI550%2B1344868036.jpg "(Smittia sp. 18ES - Finnmark550) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2012)")
![(Smittia sp. 19ES - CH-eik92) @12 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2013) NTNU Museum of Natural History and Archaeology NTNU Museum of Natural History and Archaeology](http://v3.boldsystems.org/pics/_w300/CHEIK/CH-eik92%2B1372170416.jpg "(Smittia sp. 19ES - CH-eik92) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2013)")
![(Smittia sp. 1SMG061913-THBJ040 - CBG-A13185-E12) @11 [ ] CreativeCommons - Attribution Share - Alike (2023) Centre for Biodiversity Genomics Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/GMCLB/CBG-A13185-E12%2B1695069148.jpg "(Smittia sp. 1SMG061913-THBJ040 - CBG-A13185-E12) {} [ ] CreativeCommons - Attribution Share - Alike (2023)")
![(Smittia sp. 20ES - ATNA384) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2013) NTNU Museum of Natural History and Archaeology NTNU Museum of Natural History and Archaeology](http://v3.boldsystems.org/pics/_w300/ATNA/AT384%2B1376508110.jpg "(Smittia sp. 20ES - ATNA384) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2013)")
![(Smittia sp. 21ES - DCTC_007_C8) @11 [ ] by Genome Research Ltd as Wellcome Sanger Institute Genome Research Limited operating as Wellcome Sanger Institute](http://v3.boldsystems.org/pics/_w300/DCTC/DCTC_007_C8%2B1731703324.jpg "(Smittia sp. 21ES - DCTC_007_C8) {} [ ] by")
![(Smittia sp. 22ES - BIOUG05503-E10) @14 [ ] CreativeCommons - Attribution (2013) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/SMTPC/BIOUG05503-E10%2B1368645122.jpg "(Smittia sp. 22ES - BIOUG05503-E10) {} [ ] CreativeCommons - Attribution (2013)")
![(Smittia sp. 23ES - BIOUG05503-A02) @14 [ ] CreativeCommons - Attribution (2013) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/SMTPC/BIOUG05503-A02%2B1368645866.jpg "(Smittia sp. 23ES - BIOUG05503-A02) {} [ ] CreativeCommons - Attribution (2013)")
![(Smittia sp. 24ES - BIOUG04089-A04) @14 [ ] CreativeCommons - Attribution (2013) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/GMFIC/BIOUG04089-A04%2B1363363914.jpg "(Smittia sp. 24ES - BIOUG04089-A04) {} [ ] CreativeCommons - Attribution (2013)")
![(Smittia sp. 25ES - SV548) @12 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2011) NTNU Museum of Natural History and Archaeology NTNU Museum of Natural History and Archaeology](http://v3.boldsystems.org/pics/_w300/CHRSV/SV548%2B1319822588.jpg "(Smittia sp. 25ES - SV548) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2011)")
![(Smittia sp. 26ES - SV 879) @12 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2013) NTNU Museum of Natural History and Archaeology NTNU Museum of Natural History and Archaeology](http://v3.boldsystems.org/pics/_w300/CHRSV/SV879%2B1382371534.jpg "(Smittia sp. 26ES - SV 879) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2013)")
![(Smittia sp. 27ES - SV590) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2012) NTNU Museum of Natural History and Archaeology NTNU Museum of Natural History and Archaeology](http://v3.boldsystems.org/pics/_w300/CHRSV/SV590%2B1334589846.jpg "(Smittia sp. 27ES - SV590) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2012)")
![(Smittia sp. 28ES - SV515) @12 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2011) NTNU Museum of Natural History and Archaeology NTNU Museum of Natural History and Archaeology](http://v3.boldsystems.org/pics/_w300/CHRFI/SV515%2B1304437506.jpg "(Smittia sp. 28ES - SV515) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2011)")
![(Smittia sp. 29ES - MM-CH145) @11 [ ] Creative Commons Attribution Non-Commercial Share-Alike (2020) NTNU University Museum, Department of Natural History NTNU University Museum, Department of Natural History](http://v3.boldsystems.org/pics/_w300/MMCHI/MM-CH145%2B1580158582.jpg "(Smittia sp. 29ES - MM-CH145) {} [ ] Creative Commons Attribution Non-Commercial Share-Alike (2020)")
![(Smittia sp. 2ES - SV 899) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2013) NTNU Museum of Natural History and Archaeology NTNU Museum of Natural History and Archaeology](http://v3.boldsystems.org/pics/_w300/CHRSV/SV899%2B1383330906.jpg "(Smittia sp. 2ES - SV 899) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2013)")
![(Smittia sp. 2SW - CCDB22605-D05) @12 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015) SNSB, Zoologische Staatssammlung Muenchen SNSB, Zoologische Staatssammlung Muenchen](http://v3.boldsystems.org/pics/_w300/FBCHI/CCDB22605-D05%2B1424711624.jpg "(Smittia sp. 2SW - CCDB22605-D05) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015)")
![(Smittia sp. 3ES - BIOUG17035-G10) @13 [ ] CreativeCommons - Attribution (2015) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/CNNHA/BIOUG17035-G10%2B1449260344.jpg "(Smittia sp. 3ES - BIOUG17035-G10) {} [ ] CreativeCommons - Attribution (2015)")
![(Smittia sp. 3SW - CCDB22605-E07) @11 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015) SNSB, Zoologische Staatssammlung Muenchen SNSB, Zoologische Staatssammlung Muenchen](http://v3.boldsystems.org/pics/_w300/FBCHI/CCDB22605-E07%2B1424712450.jpg "(Smittia sp. 3SW - CCDB22605-E07) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015)")
![(Smittia sp. 5ES - SV 716) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2013) NTNU Museum of Natural History and Archaeology NTNU Museum of Natural History and Archaeology](http://v3.boldsystems.org/pics/_w300/CHRSV/SV716%2B1375989966.jpg "(Smittia sp. 5ES - SV 716) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2013)")
![(Smittia sp. 6ES - BIOUG15605-D12) @13 [ ] CreativeCommons - Attribution (2015) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/GMGXX/BIOUG15605-D12%2B1427750040.jpg "(Smittia sp. 6ES - BIOUG15605-D12) {} [ ] CreativeCommons - Attribution (2015)")
![(Smittia sp. 7ES - SV597) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2012) NTNU Museum of Natural History and Archaeology NTNU Museum of Natural History and Archaeology](http://v3.boldsystems.org/pics/_w300/CHRSV/SV597%2B1334596432.jpg "(Smittia sp. 7ES - SV597) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2012)")
![(Smittia sp. 8ES - BIOUG01623-D05) @15 [ ] CreativeCommons - Attribution (2011) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/JSSEP/bioug01623-d05%2B1349355022.jpg "(Smittia sp. 8ES - BIOUG01623-D05) {} [ ] CreativeCommons - Attribution (2011)")
![(Smittia sp. 9ES - Finnmark552) @11 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2012) NTNU Museum of Natural History and Archaeology NTNU Museum of Natural History and Archaeology](http://v3.boldsystems.org/pics/_w300/CHRFI/FI552%2B1344868270.jpg "(Smittia sp. 9ES - Finnmark552) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2012)")
![(Smittia sp. ES11 - BIOUG03578-A03) @13 [ ] CreativeCommons - Attribution (2013) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/CNEID/BIOUG03578-A03%2B1377093312.jpg "(Smittia sp. ES11 - BIOUG03578-A03) {} [ ] CreativeCommons - Attribution (2013)")
![(Smittia sp. ES12 - BIOUG16790-H03) @15 [ ] CreativeCommons - Attribution (2015) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/CNIVB/BIOUG16790-H03%2B1424275132.jpg "(Smittia sp. ES12 - BIOUG16790-H03) {} [ ] CreativeCommons - Attribution (2015)")
![(Smittia stercoraria - ZMUO.025335) @12 [ ] CreativeCommons - Attribution Non-Commercial (2015) Marko Mutanen University of Oulu](http://v3.boldsystems.org/pics/_w300/CHIFI/ZMUO.025335%2B1450728926.JPG "(Smittia stercoraria - ZMUO.025335) {} [ ] CreativeCommons - Attribution Non-Commercial (2015)")
![(Smittia terrestris - ZFMK-TIS-2603350) @11 [ ] CreativeCommons Attribution Share-Alike (by-sa) 818 (2017) Unspecified Zoologisches Forschungsmuseum Alexander Koenig](http://v3.boldsystems.org/pics/_w300/GDIP/ZFMK_2603350%2B1501598768.jpg "(Smittia terrestris - ZFMK-TIS-2603350) {} [ ] CreativeCommons Attribution Share-Alike (by-sa) 818 (2017)")
