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Identification
Workbench
Resources
Latest Version
Pentodontini {tribe} -
Arthropoda
;
Insecta
;
Coleoptera
;
Scarabaeidae
;
Dynastinae
;
Print
For the latest taxonomy data, visit
boldsystems.org
.
Search Taxonomy
Sub-taxa
Genera (34)
Alissonotum [26]
Anomalomorpha [6]
Anoplognatho [2]
Aphonus [18]
Barutus [5]
Bothynus [14]
Calicnemis [71]
Carneodon [3]
Cheiroplatys [2]
Coscinocephalus [7]
Dasygnathus [24]
Diloboderus [41]
Euetheola [38]
Eutyctus [2]
Heteronychus [57]
Mellissius [1]
Metanastes [5]
Neocorynophyllus [1]
Novapus [1]
Orizabus [14]
Oxygrylius [13]
Oxyligyrus [5]
Parapucaya [8]
Pentodon [44]
Pericoptus [314]
Phyllognathus [16]
Pimelopus [3]
Pucaya [10]
Pycnoschema [1]
Semanopterus [5]
Temnorhynchus [7]
Thronistes [5]
Tomarus [150]
Trissodon [1]
Taxon Description (Wikipedia)
full article at Wikipedia
BOLD Stats
Specimen Records:
923
Public Records:
401
Specimens with Sequences:
801
Public Species:
75
Specimens with Barcodes:
688
Public BINs:
0
Species:
92
Species With Barcodes:
65
Species List - Progress
Access Published & Released Data
Contributors (Specimens & Sequencing)
Specimen Depositories:
Sequencing Labs:
Imagery
images representing subtaxa of Pentodontini
(
Tomarus
)
Sample ID:
CCDB-23522-G09
License:
CreativeCommons - Attribution (2015)
License Holder:
CBG Photography Group, Centre for Biodiversity Genomics
Collection Sites
Collected from 25 countries.
Top 20:
Show All Countries
United States
91
Argentina
16
Georgia
4
Costa Rica
76
Spain
15
Mexico
3
Australia
60
France
14
Austria
3
Peru
23
Ecuador
12
New Zealand
3
Italy
20
Panama
6
Pakistan
3
Canada
18
Colombia
6
Sri Lanka
2
South Africa
18
India
5
![(Anomalomorpha - ww03705) @14 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2017) Andrew Mitchell Australian Museum](http://v3.boldsystems.org/pics/_w300/SCARB/ww03705%2B1491852362.jpg "(Anomalomorpha - ww03705) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2017)")
![(Anoplognatho - CNC COLEO 00124683) @13 [ ] CreativeCommons - Attribution (2011) CNC/CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/CNCCC/CNC_COLEO_00124683%2B1309885798.JPG "(Anoplognatho - CNC COLEO 00124683) {} [ ] CreativeCommons - Attribution (2011)")
![(Aphonus - CNC COLEO 00124691) @15 [ ] CreativeCommons - Attribution (2011) CNC/CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/CNCCC/CNC_COLEO_00124691%2B1309885890.JPG "(Aphonus - CNC COLEO 00124691) {} [ ] CreativeCommons - Attribution (2011)")
![(Barutus - INB0003044280) @15 [ ] Copyright (2010) A. Solis Instituto Nacional de Biodiversidad](http://v3.boldsystems.org/pics/_w300/ASSCR/INB0003044280%2B1286913972.jpg "(Barutus - INB0003044280) {} [ ] Copyright (2010)")
![(Bothynus - INBIOCRI002328754) @14 [ ] Copyright (2010) A. Solis Instituto Nacional de Biodiversidad](http://v3.boldsystems.org/pics/_w300/ASSCR/INBIOCRI002328754%2B1286913974.jpg "(Bothynus - INBIOCRI002328754) {} [ ] Copyright (2010)")
![(Calicnemis - CTBB-0837) @15 [ ] Copyright (2013) Thierry Bouyer Research Collection of Thierry Bouyer](http://v3.boldsystems.org/pics/_w300/TBMOT/CTBB%5B837%5D%2B1383949936.jpg "(Calicnemis - CTBB-0837) {} [ ] Copyright (2013)")
![(Coscinocephalus - CCDB-23522-F03) @14 [ ] CreativeCommons - Attribution (2015) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/LYMAB/CCDB-23522-F03%2B1444413512.JPG "(Coscinocephalus - CCDB-23522-F03) {} [ ] CreativeCommons - Attribution (2015)")
![(Dasygnathus - ww04331) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2017) Andrew Mitchell Australian Museum](http://v3.boldsystems.org/pics/_w300/SCARB/ww04331%2B1216927254.jpg "(Dasygnathus - ww04331) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2017)")
![(Diloboderus - MACN ASI MKC 0059) @11 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2012) Paula Florencia Zermoglio, Jose; Emilio Crespo Museo Argentino de Ciencias Naturales](http://v3.boldsystems.org/pics/_w300/MKCSC/MACN_ASI_MKC_0059_2%2B1346524458.JPG "(Diloboderus - MACN ASI MKC 0059) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2012)")
![(Euetheola - BIOUG02025-E08) @14 [ ] CreativeCommons - Attribution (2012) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/BBCCA/IMG_9792%2B1333984188.JPG "(Euetheola - BIOUG02025-E08) {} [ ] CreativeCommons - Attribution (2012)")
![(Eutyctus - BC-TB6341) @11 [ ] Copyright (2010) Thierry Bouyer Unspecified](http://v3.boldsystems.org/pics/_w300/CERAF/BC-TB%5B6341%5D%2B1286380362.jpg "(Eutyctus - BC-TB6341) {} [ ] Copyright (2010)")
![(Heteronychus - BIOUG02563-C11) @15 [ ] CreativeCommons - Attribution (2017) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/COAS/BIOUG02563-C11%2B1491491254.jpg "(Heteronychus - BIOUG02563-C11) {} [ ] CreativeCommons - Attribution (2017)")
![(Mellissius - NHMUK014666312) @11 [ ] CreativeCommons Attribution (by) Trustees of the Natural History Museum, London Natural History Museum, London](http://v3.boldsystems.org/pics/_w300/HELEN/014666312_Dorsal%2B1675739294.jpg "(Mellissius - NHMUK014666312) {} [ ] CreativeCommons Attribution (by)")
![(Metanastes - ww04285) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2017) Andrew Mitchell Australian Museum](http://v3.boldsystems.org/pics/_w300/SCARB/ww04285%2B1216993232.jpg "(Metanastes - ww04285) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2017)")
![(Orizabus - BIOUG02023-F01) @15 [ ] CreativeCommons - Attribution (2012) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/BBCCA/IMG_9544%2B1333636474.jpg "(Orizabus - BIOUG02023-F01) {} [ ] CreativeCommons - Attribution (2012)")
![(Oxygrylius - BIOUG02025-G01) @14 [ ] CreativeCommons - Attribution (2012) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/BBCCA/IMG_9823%2B1333983536.jpg "(Oxygrylius - BIOUG02025-G01) {} [ ] CreativeCommons - Attribution (2012)")
![(Oxyligyrus - QCAZI_981) @11 [ ] CC-BY-NC-ND (2024) QCAZ Pontificia Universidad Católica del Ecuador](http://v3.boldsystems.org/pics/_w300/SCAEC/QCAZI_981%2B1744057484.jpg "(Oxyligyrus - QCAZI_981) {} [ ] CC-BY-NC-ND (2024)")
![(Parapucaya - INBIOCRI001632291) @13 [ ] Copyright (2010) A. Solis Instituto Nacional de Biodiversidad](http://v3.boldsystems.org/pics/_w300/ASSCR/INBIOCRI001632291%2B1331843656.jpg "(Parapucaya - INBIOCRI001632291) {} [ ] Copyright (2010)")
![(Pentodon - GBOL_Col_FK_7377) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015) SNSB, Zoologische Staatssammlung Muenchen SNSB, Zoologische Staatssammlung Muenchen](http://v3.boldsystems.org/pics/_w300/GBCOU/GBOL_Col_FK_7377%2B1378920068.jpg "(Pentodon - GBOL_Col_FK_7377) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015)")
![(Pericoptus - NZ750_2023) @11 [ ] Unspecified (default): All Rights Reserved Unspecified Unspecified](http://v3.boldsystems.org/pics/_w300/LMOUY/NZ750_Dorsal%2B1681921672.jpg "(Pericoptus - NZ750_2023) {} [ ] Unspecified (default): All Rights Reserved")
![(Pimelopus - ww04284) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2017) Andrew Mitchell Australian Museum](http://v3.boldsystems.org/pics/_w300/SCARB/ww04284%2B1216926640.jpg "(Pimelopus - ww04284) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2017)")
![(Pucaya - INB0003737688) @14 [ ] Copyright (2010) A. Solis Instituto Nacional de Biodiversidad](http://v3.boldsystems.org/pics/_w300/ASSCR/INB0003737688%2B1286913998.jpg "(Pucaya - INB0003737688) {} [ ] Copyright (2010)")
![(Semanopterus - BIOUG02565-B03) @14 [ ] CreativeCommons - Attribution (2013) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/COAS/BIOUG02565-B03%2B1362430134.jpg "(Semanopterus - BIOUG02565-B03) {} [ ] CreativeCommons - Attribution (2013)")
![(Temnorhynchus - DNA_T533) @11 [ ] © (2024) Thomas Schmitzer Julius-Maximilians-Universität Würzburg](http://v3.boldsystems.org/pics/_w300/ANDIV/DNA_T533%2B1697219888.jpg "(Temnorhynchus - DNA_T533) {} [ ] © (2024)")
![(Tomarus - CCDB-23522-G09) @15 [ ] CreativeCommons - Attribution (2015) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/LYMAB/CCDB-23522-G09%2B1444415236.JPG "(Tomarus - CCDB-23522-G09) {} [ ] CreativeCommons - Attribution (2015)")
