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Latest Version
Harpalinae {subfamily} -
Arthropoda
;
Insecta
;
Coleoptera
;
Carabidae
;
Print
For the latest taxonomy data, visit
boldsystems.org
.
Search Taxonomy
Sub-taxa
Tribes (37)
Abacetini [76]
Anthiini [20]
Atranini [8]
Calophaenini [8]
Chlaeniini [1296]
Cnemalobini [8]
Collyridini [27]
Corsyrini [8]
Ctenodactylini [21]
Cyclosomini [226]
Dercylini [29]
Drimostomatini [2]
Dryptini [30]
Galeritini [310]
Harpalini [10937]
Helluonini [235]
Hexagoniini [4]
Lachnophorini [56]
Lebiini [3648]
Licinini [1357]
Masoreini [27]
Metiini [52]
Morionini [22]
Odacanthini [116]
Oodini [208]
Orthogoniini [8]
Panagaeini [110]
Peleciini [7]
Pentagonicini [254]
Perigonini [60]
Physocrotaphini [2]
Platynini [4587]
Pseudomorphini [27]
Pterostichini [10429]
Sphodrini [2631]
Zabrini [4416]
Zuphiini [80]
Genera (1)
Selenophori [1]
Taxon Description (Wikipedia)
full article at Wikipedia
BOLD Stats
Specimen Records:
44,450
Public Records:
35,221
Specimens with Sequences:
39,075
Public Species:
1,876
Specimens with Barcodes:
35,691
Public BINs:
1,636
Species:
2,148
Species With Barcodes:
1,575
Species List - Progress
Access Published & Released Data
Contributors (Specimens & Sequencing)
Specimen Depositories:
Sequencing Labs:
Imagery
images representing subtaxa of Harpalinae
(
Zuphiini
)
Sample ID:
CNC COLEO 00119546
License:
CreativeCommons - Attribution (2011)
License Holder:
CNC/CBG Photography Group, Centre for Biodiversity Genomics
Collection Sites
Collected from 75 countries.
Top 20:
Show All Countries
United States
19904
Norway
266
Colombia
135
Canada
5428
Netherlands
210
Ecuador
125
Germany
4289
Australia
183
Croatia
119
Costa Rica
2363
United Kingdom
180
Finland
107
Austria
2096
Peru
163
South Africa
102
Ghana
515
Pakistan
157
Greece
64
France
349
Bulgaria
146
![(Abacetini - CCDB-21417-D05) @14 [ ] CreativeCommons - Attribution (2014) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/LYMAA/ccdb-21417-d05%2B1401386178.jpg "(Abacetini - CCDB-21417-D05) {} [ ] CreativeCommons - Attribution (2014)")
![(Anthiini - NIBGE COL-01379) @14 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2011) Muhammad Ashfaq National Institute for Biotechnology and Genetic Engineering Faisalabad Pakistan](http://v3.boldsystems.org/pics/_w300/MACOL/NIBGE_COL-01379%2B1311036506.JPG "(Anthiini - NIBGE COL-01379) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2011)")
![(Atranini - CCDB-21331-E10) @15 [ ] CreativeCommons - Attribution (2014) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/LYMAA/ccdb-21331-e10%2B1397228426.jpg "(Atranini - CCDB-21331-E10) {} [ ] CreativeCommons - Attribution (2014)")
![(Calophaenini - BC-GF2024-0711) @11 [ ] Creative Commons (2024) rodolphe rougerie Museum national d'histoire naturelle](http://v3.boldsystems.org/pics/_w300/FGBEL/GF_0711%2B315525600.JPG "(Calophaenini - BC-GF2024-0711) {} [ ] Creative Commons (2024)")
![(Chlaeniini - BCZSM_COLA_01547) @16 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2012) SNSB, Zoologische Staatssammlung Muenchen SNSB, Zoologische Staatssammlung Muenchen](http://v3.boldsystems.org/pics/_w300/FBCOH/BCZSM_COLA_01547%2B1350412280.JPG "(Chlaeniini - BCZSM_COLA_01547) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2012)")
![(Cnemalobini - CCDB-30443-A02) @11 [ ] CreativeCommons - Attribution (2018) Smithsonian Institution Smithsonian Institution](http://v3.boldsystems.org/pics/_w300/SICOA/CCDB-30443-A02_Dorsal%2B1540910400.jpg "(Cnemalobini - CCDB-30443-A02) {} [ ] CreativeCommons - Attribution (2018)")
![(Collyridini - NIBGE COL-01149) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2011) Muhammad Ashfaq National Institute for Biotechnology and Genetic Engineering Faisalabad Pakistan](http://v3.boldsystems.org/pics/_w300/MACOL/NIBGE_COL-1149%2B1307714192.jpg "(Collyridini - NIBGE COL-01149) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2011)")
![(Corsyrini - CCDB-30443-C04) @11 [ ] CreativeCommons - Attribution (2018) Smithsonian Institution Smithsonian Institution](http://v3.boldsystems.org/pics/_w300/SICOA/CCDB-30443-C04_Dorsal%2B1542137280.jpg "(Corsyrini - CCDB-30443-C04) {} [ ] CreativeCommons - Attribution (2018)")
![(Ctenodactylini - BIOUG02279-E10) @14 [ ] CreativeCommons - Attribution (2012) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/BBCCA/bioug02279-e10%2B1354034360.JPG "(Ctenodactylini - BIOUG02279-E10) {} [ ] CreativeCommons - Attribution (2012)")
![(Cyclosomini - BIOUG02279-D11) @14 [ ] CreativeCommons - Attribution (2012) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/BBCCA/bioug02279-d11%2B1354034264.JPG "(Cyclosomini - BIOUG02279-D11) {} [ ] CreativeCommons - Attribution (2012)")
![(Dercylini - CCDB-30447-A09) @11 [ ] CreativeCommons - Attribution (2018) Smithsonian Institution Smithsonian Institution](http://v3.boldsystems.org/pics/_w300/SICOA/CCDB-30447-A09_Lateral%2B1536860040.jpg "(Dercylini - CCDB-30447-A09) {} [ ] CreativeCommons - Attribution (2018)")
![(Drimostomatini - CCDB-30450-G08) @11 [ ] CreativeCommons - Attribution (2018) Smithsonian Institution Smithsonian Institution](http://v3.boldsystems.org/pics/_w300/SICOA/CCDB-30450-G08_Dorsal%2B1539098494.jpg "(Drimostomatini - CCDB-30450-G08) {} [ ] CreativeCommons - Attribution (2018)")
![(Dryptini - BC ZSM COL 01911) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2010) Stefan Schmidt SNSB, Zoologische Staatssammlung Muenchen](http://v3.boldsystems.org/pics/_w300/FBCOC/BCZSMCOL01911%2B1284714370.JPG "(Dryptini - BC ZSM COL 01911) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2010)")
![(Galeritini - CCDB-21430-D07) @15 [ ] CreativeCommons - Attribution (2014) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/LYMAA/CCDB-21430-D07%2B1406035248.JPG "(Galeritini - CCDB-21430-D07) {} [ ] CreativeCommons - Attribution (2014)")
![(Harpalini - 09BBCOL-0669) @16 [ ] CreativeCommons - Attribution (2009) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/USCOL/IMG_8614%2B1250514704.JPG "(Harpalini - 09BBCOL-0669) {} [ ] CreativeCommons - Attribution (2009)")
![(Helluonini - 10BBCOL-0116) @15 [ ] CreativeCommons - Attribution (2010) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/USCOC/10bbcol-0116%2B1279553772.jpg "(Helluonini - 10BBCOL-0116) {} [ ] CreativeCommons - Attribution (2010)")
![(Hexagoniini - CCDB-30450-A05) @11 [ ] CreativeCommons - Attribution (2018) Smithsonian Institution Smithsonian Institution](http://v3.boldsystems.org/pics/_w300/SICOA/CCDB-30450-A05_Dorsal%2B1537972320.jpg "(Hexagoniini - CCDB-30450-A05) {} [ ] CreativeCommons - Attribution (2018)")
![(Lachnophorini - NEONTcarabid5581) @15 [ ] Copyright (2012) Barton, M National Ecological Observatory Network (NEON) http://www.neoninc.org/content/copyright](http://v3.boldsystems.org/pics/_w300/NEONT/carabid5581%2B1369403022.jpg "(Lachnophorini - NEONTcarabid5581) {} [ ] Copyright (2012)")
![(Lebiini - BIOUG00972-B09) @16 [ ] CreativeCommons - Attribution (2011) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/JSCOL/bioug00972-b09%2B1311770604.jpg "(Lebiini - BIOUG00972-B09) {} [ ] CreativeCommons - Attribution (2011)")
![(Licinini - CCDB-21431-C12) @15 [ ] CreativeCommons - Attribution (2014) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/LYMAA/ccdb-21431-c12%2B1401471714.jpg "(Licinini - CCDB-21431-C12) {} [ ] CreativeCommons - Attribution (2014)")
![(Masoreini - ZFMK_COL_2010_660) @13 [ ] CreativeCommons - Attribution (2011) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/EUCAR/zfmk_col_2010_660%2B1311867238.jpg "(Masoreini - ZFMK_COL_2010_660) {} [ ] CreativeCommons - Attribution (2011)")
![(Metiini - CCDB-30450-B12) @11 [ ] CreativeCommons - Attribution (2018) Smithsonian Institution Smithsonian Institution](http://v3.boldsystems.org/pics/_w300/SICOA/CCDB-30450-B12_Dorsal%2B1537988400.jpg "(Metiini - CCDB-30450-B12) {} [ ] CreativeCommons - Attribution (2018)")
![(Morionini - CCDB-21406-F04) @14 [ ] CreativeCommons - Attribution (2014) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/LYMAA/ccdb-21406-f04%2B1397593066.jpg "(Morionini - CCDB-21406-F04) {} [ ] CreativeCommons - Attribution (2014)")
![(Odacanthini - ZFMK_COL_2011_227) @14 [ ] CreativeCommons - Attribution (2011) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/EUCAR/ccdb-12751-f03%2B1311283620.jpg "(Odacanthini - ZFMK_COL_2011_227) {} [ ] CreativeCommons - Attribution (2011)")
![(Oodini - CCDB-21431-H02) @15 [ ] CreativeCommons - Attribution (2014) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/LYMAA/ccdb-21431-h02%2B1401713120.jpg "(Oodini - CCDB-21431-H02) {} [ ] CreativeCommons - Attribution (2014)")
![(Orthogoniini - CCDB-30450-D09) @11 [ ] CreativeCommons - Attribution (2018) Smithsonian Institution Smithsonian Institution](http://v3.boldsystems.org/pics/_w300/SICOA/CCDB-30450-D09_Lateral%2B1538060220.jpg "(Orthogoniini - CCDB-30450-D09) {} [ ] CreativeCommons - Attribution (2018)")
![(Panagaeini - ZMUO.029353) @15 [ ] CreativeCommons - Attribution Non-Commercial (2017) Marko Mutanen University of Oulu](http://v3.boldsystems.org/pics/_w300/COLFH/ZMUO.029353%2B1496100632.JPG "(Panagaeini - ZMUO.029353) {} [ ] CreativeCommons - Attribution Non-Commercial (2017)")
![(Peleciini - CCDB-30450-D12) @11 [ ] CreativeCommons - Attribution (2018) Smithsonian Institution Smithsonian Institution](http://v3.boldsystems.org/pics/_w300/SICOA/CCDB-30450-D12_Dorsal%2B1538061180.jpg "(Peleciini - CCDB-30450-D12) {} [ ] CreativeCommons - Attribution (2018)")
![(Pentagonicini - BIOUG22791-G09) @14 [ ] CreativeCommons - Attribution (2015) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/GMACF/BIOUG22791-G09%2B1442345580.jpg "(Pentagonicini - BIOUG22791-G09) {} [ ] CreativeCommons - Attribution (2015)")
![(Perigonini - RMNH.INS.535232) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2013) Unspecified Naturalis Biodiversity Center](http://v3.boldsystems.org/pics/_w300/NLCOA/RMNH.INS.535232_dor%2B1445540330.jpg "(Perigonini - RMNH.INS.535232) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2013)")
![(Physocrotaphini - CCDB-30450-E05) @11 [ ] CreativeCommons - Attribution (2018) Smithsonian Institution Smithsonian Institution](http://v3.boldsystems.org/pics/_w300/SICOA/CCDB-30450-E05_Dorsal%2B1538068800.jpg "(Physocrotaphini - CCDB-30450-E05) {} [ ] CreativeCommons - Attribution (2018)")
![(Platynini - CCDB-21333-F06) @15 [ ] CreativeCommons - Attribution (2014) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/LYMAA/ccdb-21333-f06%2B1397063014.jpg "(Platynini - CCDB-21333-F06) {} [ ] CreativeCommons - Attribution (2014)")
![(Pseudomorphini - CCDB-30448-A04) @11 [ ] CreativeCommons - Attribution (2018) Smithsonian Institution Smithsonian Institution](http://v3.boldsystems.org/pics/_w300/SICOA/CCDB-30448-A04_Dorsal%2B1537790340.jpg "(Pseudomorphini - CCDB-30448-A04) {} [ ] CreativeCommons - Attribution (2018)")
![(Pterostichini - CCDB-21431-C04) @15 [ ] CreativeCommons - Attribution (2014) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/LYMAA/ccdb-21431-c04%2B1401471530.jpg "(Pterostichini - CCDB-21431-C04) {} [ ] CreativeCommons - Attribution (2014)")
![(Sphodrini - CNC COLEO 00118598) @16 [ ] CreativeCommons - Attribution (2011) CNC/CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/CNCCB/CNC_COLEO_00118598%2B1307982920.JPG "(Sphodrini - CNC COLEO 00118598) {} [ ] CreativeCommons - Attribution (2011)")
![(Zabrini - BIOUG14142-E05) @15 [ ] CreativeCommons - Attribution (2015) CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/SSROB/BIOUG14142-E05%2B1422649230.jpg "(Zabrini - BIOUG14142-E05) {} [ ] CreativeCommons - Attribution (2015)")
![(Zuphiini - CNC COLEO 00119546) @13 [ ] CreativeCommons - Attribution (2011) CNC/CBG Photography Group Centre for Biodiversity Genomics](http://v3.boldsystems.org/pics/_w300/CNCCB/CNC_COLEO_00119546%2B1308231618.JPG "(Zuphiini - CNC COLEO 00119546) {} [ ] CreativeCommons - Attribution (2011)")
